Wendell R. HaagResearch Fisheries Biologist
3761 Georgetown Road
Frankfort, KY 40601
I conduct research on the ecology of aquatic organisms and how this information can be used to develop effective management and conservation strategies for streams. Most of my current research is focused on understanding causes of freshwater mussel declines throughout the U.S. and much of the world. Mussels are important because, as filter feeders, they have a profound influence on aquatic ecosystem function and health, and a large percentage of North American species are in danger of extinction. My research on mussel declines includes a wide variety of field and laboratory approaches conducted with a broad network of partners. I have also conducted extensive research on freshwater fishes and have experience with aquatic insects, crayfishes, snails, reptiles and amphibians, and many other groups.
- Ph.D. in Biology, 2002
- University of Mississippi
- M.S. in Zoology, 1991
- The Ohio State University
- B.S. in Biology, 1988
- Eastern Kentucky University
- Haag, Wendell R.; Stoeckel, James A. 2021. Demographic characteristics of young-of-year freshwater mussel populations in ponds.
- Douda, Karel ; Haag, Wendell R.; Escobar-Calderón, Felipe ; Vodáková, Barbora ; Reichard, Martin ; Chen, Xiubao ; McGregor, Monte ; Yang, Jian ; Lopes-Lima, Manuel . 2021. Effects of in vitro metamorphosis on survival, growth, and reproductive success of freshwater mussels.
- Mayfield, Albert (Bud); Seybold, Steven; Haag, Wendell; Johnson, Tracy; Kerns, Becky; Kilgo, John; Larkin, Daniel; Lucardi, Rima; Moltzan, Bruce; Pearson, Dean; Rothlisberger, John; Schardt, John; Schwartz, Michael; Young, Michael. 2021. Impacts of invasive species in terrestrial and aquatic systems in the United States.
- Haag, Wendell R.; Culp, Jacob ; Drayer, Andrea N.; McGregor, Monte A.; White, Drew E. J.; Price, Steven J. 2020. Abundance of an invasive bivalve, Corbicula fluminea , is negatively related to growth of freshwater mussels in the wild.
- Atkinson, Carla L.; Parr, Thomas B.; Ee, Brian C. van; Knapp, Daniel D.; Winebarger, Monica ; Madoni, Kyle J.; Haag, Wendell R. 2020. Length-mass equations for freshwater unionid mussel assemblages: Implications for estimating ecosystem function.
- Strayer, David L.; Geist, Juergen ; Haag, Wendell R.; Jackson, John K.; Newbold, J. Denis. 2019. Essay: Making the most of recent advances in freshwater mussel propagation and restoration.
- Haag, Wendell R.; Culp, J. Jacob; McGregor, Monte A.; Bringolf, Robert ; Stoeckel, James A. 2019. Growth and survival of juvenile freshwater mussels in streams: Implications for understanding enigmatic mussel declines.
- Haag, Wendell R. 2019. Reassessing enigmatic mussel declines in the United States.
- Hart, Michael A.; Haag, Wendell R.; Bringolf, Robert ; Stoeckel, James A. 2018. Novel technique to identify large river host fish for freshwater mussel propagation and conservation.
- Fritts, Andrea K.; Fritts, Mark W.; Haag, Wendell R.; DeBoer, Jason A.; Casper, Andrew F. 2017. Freshwater mussel shells (Unionidae) chronicle changes in a North American river over the past 1000 years.
- Dolloff, Andy; Roghair, Craig; Krause, Colin; Moran, John; Cochran, Allison; Warren, Mel; Adams, Susan; Haag, Wendell. 2016. Spatio-temporal variation in distribution of aquatic species and their habitats in a reservoir transition zone.
- Haag, Wendell R.; Stoeckel, James A. 2015. The role of host abundance in regulating populations of freshwater mussels with parasitic larvae.
- Klunzinger, Michael W.; Beatty, Stephen J.; Morgan, David L.; Lymbery, Alan J.; Haag, Wendell R. 2014. Age and growth in the Australian freshwater mussel, Westralunio carteri , with an evaluation of the fluorochrome calcein for validating the assumption of annulus formation.
- Haag, Wendell R.; Williams, James D. 2014. Biodiversity on the brink: an assessment of conservation strategies for North American freshwater mussels.
- Mosley, Tyler L.; Haag, Wendell R.; Stoeckel, James A. 2014. Egg fertilisation in a freshwater mussel: effects of distance, flow and male density.
- Gascho-Landis, Andrew M.; Haag, Wendell R.; Stoeckel, James A. 2013. High suspended solids as a factor in reproductive failure of a freshwater mussel.
- Haag, Wendell R.; Distefano, Robert J.; Fennessy, Siobhan; Marshall, Brett D. 2013. Invertebrates and Plants.
- Haag, Wendell R.; Berg, D.J.; Galbraith, H.: Spooner, D..; Strayer, D.L. 2013. Professional awards (Freshwater Mollusk Conservation Society).
- Haag, Wendell R. 2013. The role of fecundity and reproductive effort in defining life history strategies of North American freshwater mussels.
- Gascho-Landis, Andrew M.; Mosley, Tyler L.; Haag, Wendell R.; Stoeckel, James A. 2012. Effects of temperature and photoperiod on lure display and glochidial release in a freshwater mussel.
- Haag, Wendell R. 2012. North American freshwater mussels: natural history, ecology, and conservation.
- Culp, Jacob J.; Haag, Wendell R.; Arrington, D. Albrey; Kennedy, Thomas B. 2011. Seasonal and species-specific patterns in abundance of freshwater mussel glochidia in stream drift.
- Haag, Wendell R. 2010. A hierarchical classification of freshwater mussel diversity in North America.
- Haag, Wendell R.; Warren, Melvin L. Jr. 2010. Diversity, abundance, and size structure of bivalve assemblages in the Sipsey River, Alabama.
- Haag, Wendell R.; Rypel, Andrew L. 2010. Growth and longevity in freshwater mussels: evolutionary and conservation implications.
- Warren, Melvin L., Jr..; Sheldon, A.L.; Haag, W.R. 2009. Constructed microhabitat bundles for sampling fishes and crayfishes in coastal plain streams.
- Haag, Wendell R. 2009. Extreme longevity in freshwater mussels revisited: sources of bias in age estimates derived from mark-recapture experiments.
- Haag, Wendell R. 2009. Past and future patterns of freshwater mussel extinctions in North America during the Holocene.
- Rypel, Andrew L.; Haag, Wendell R.; Findlay, Robert H. 2009. Pervasive hydrologic effects on freshwater mussels and riparian trees in southeastern floodplain ecosystems.
- Rypel, Andrew L.; Haag, Wendell R.; Findlay, Robert H. 2009. Validation of annual growth rings in freshwater mussel shells using cross dating .Can.
- Barnhart, Christopher M.; Haag, Wendell R.; Roston, William N. 2008. Adaptations to host infection and larval parasitism in Unionoida.
- Haag, Wendell R.; Commens-Carson, Amy M. 2008. Testing the assumption of annual shell ring deposition in freshwater mussels.
- Haag, Wendell R.; Warren, Melvin L. 2007. Freshwater mussel assemblage structure in a regulated river in the Lower Mississippi river Alluvial Basin, USA.
- Haag, Wendell R.; Warren, Melvin L., Jr. 2006. Seasonal feeding specialization on snails by river darters (Percina shumardi) with a review of snail feeding by other darter species.
- Peacock, Evan; Haag, Wendell, R.; Warren, Melvin, L. 2005. Prehistoric decline in freshwater mussels coincident with the advent of maize agriculture.
- Warren, Melvin L., Jr..; Haag, Wendell R. 2005. Spatio-temporal patterns of the decline of fresh water mussels in the Little South Fork Cumberland River,USA.
- Kennedy, Thomas B.; Haag, Wendell R. 2005. Using morphometrics to identify glochidia from a diverse freshwater mussel community.
- Strayer, David L.; Downing, John A.; Haag, Wendell R.; King, Timothy L.; Layzer, James B.; Newton, Teresa J.; Nichols, S. Jerrine. 2004. Changing perspectives on pearly mussels, North America's most imperiled animals.
- Warren, Melvin L., Jr..; Haag, Wendell R.; Commens, Amy M. 2004. Observations of watersnake (Nerodia, Colubridae predation on Darters (Percidae).
- Adams, Susan B.; Warren, Melvin L., Jr..; Haag, Wendell R. 2004. Spatial and temporal patterns in fish assemblages of upper coastal plain streams, Mississippi, USA.
- Haag, Wendell R.; Warren, Melvin L., Jr. 2003. Host fishes and infection strategies of freshwater mussels in large Mobile Basin streams, USA.
- Haag, Wendell R.; Staton, J. Leann. 2003. Variation in fecundity and other reproductive traits in freshwater mussels.
- Haag, Wendell R.; Warren, Melvin L., Jr..; Wright, Keith; Shaffer, Larry. 2002. Occurrence of the Rayed Creekshell, Anodontoides Radzatus, in the Mississippi River Basin: Implications For Conservation and Biogeography.
- Haag, Wendell R.; Warren, Melvin L., Jr. 2000. Effects of light and presence of fish on lure display and larval release behaviours in two species of freshwater mussels.
- Warren, Melvin L., Jr..; Haag, Wendell R.; Adams, Susan B. 2000. Forest linkages to diversity and abundancein lowland stream fish communities.
- Haag, Wendell R.; Warren, Melvin L., Jr..; Shillingsford, Mahala. 1999. Host fishes and host-attracting behavior of Lampsilis altilis and Villosa vibex (Bivalvia: Unionidae).
- Haag, Wendell R.; Warren, Melvin L., Jr. 1999. Mantle displays of freshwater mussels elicit attacks from fish.
- Warren, Melvin L., Jr..; Haag, Wendell R.; Burr, Brooks M. 1999. Status of the Mussel Resource in Little South Fork Cumberland River.
- Haag, Wendell R.; Warren, Melvin L., Jr. 1998. Freshwater mussels of the Delta National Forest, Mississippi Final Report.
- Haag, Wendell R.; Warren, Melvin L., Jr. 1998. Role of ecological factors and reproductive strategies in structuring freshwater mussel communities.
- Warren, Melvin L., Jr..; Angermeier, Paul L.; Burr, Brooks M.; Haag, Wendell R. 1997. Decline of a diverse fish fauna: patterns of imperilment and protection in the Southeastern United States.
- Haag, Wendell R.; Warren, Melvin L., Jr. 1997. Host fishes and reproductive biology of 6 freshwater mussel species from the Mobile Basin, USA.
- Johnston, Carol E.; Haag, Wendell R. 1996. Life History Of The Yazoo Darter (Percidae: Etheostoma Raneyi), A Species Endemic To North-Central Mississippi.
- Haag, Wendell; Butler, Robert S.; Hartfield, Paul D. 1995. An extraordinary reproductive strategy in freshwater bivalves: prey mimicry to facilitate larval dispersal.
- Haag, Wendell R.; Warren, Melvin L., Jr. 1995. Current Distributional Information on Freshwater Mussels (family Unionidae) in Mississippi National Forests.
- Berg, David J.; Haag, Wendell R.; Guttman, Sheldon I.; Sickel, James B. 1995. Mantle biopsy: a technique for nondestructive tissue-sampling of freshwater mussels.
- Haag, Wendell R.; Garton, David W. 1995. Variation in Genotype Frequencies During the Life History of the Bivalve, Dreissena Polymorpha.
- Assessment of conservation strategies for North American freshwater mussels. (2014)
- North America has the most diverse freshwater mussel fauna on Earth, but over half of the 300 native species are in danger of extinction. Extensive conservation efforts in the last 30 years have addressed this extinction crisis. A Forest Service scientist critically assessed the effectiveness of those efforts and outlined ways that conservation can be more effective in the future.